Working Papers

Ellingsen, Herrmann, MA Nowak, DG Rand, and CE Tarnita. “Civic Capital in Two Cultures: The Nature of Cooperation in Romania and USA..” (2013): n. pag. Print.


Martinez-Garcia, Ricardo, CE Tarnita, and Juan A. Bonachela. “Spatial Patterns in Ecological Systems: From Microbial Colonies to Landscapes.” Emerging Topics in Life Sciences (2022): n. pag. Print.
Staps, Merlijn, and CE Tarnita. “When Being Flexible Matters: Ecological Underpinnings for the Evolution of Collective Flexibility and Task Allocation.” PNAS 119.18 (2022): e2116066119. Print.
Staps, Merlijn, Jordi Gestel, and CE Tarnita. “Life Cycles As a Central Organizing Theme for Studying Multicellularity.” The Evolution of Multicellularity. Boca Raton: CRC Press, 2022. Print.
Strogatz, Steven, Sara Walker, Julia Yeomans, CE Tarnita, Elsa Arcaute, Manlio De Domenico, Oriol Artime, and Kwang-Il Goh. “Fifty Years of ‘More Is different’.” Nature Reviews Physics (2022): n. pag.


Castillo Vardaro, Jessica, Juan Bonachela, Christopher Baker, Malin Pinsky, Daniel Doak, Robert Pringle, and Corina Tarnita. “Resource Availability and Heterogeneity Shape the Self-Organization of Regular Spatial Patterning.” Ecology Letters (2021): n. pag.
Coverdale, Tyler, Ryan O’Connell, Matthew Hutchinson, Amanda Savagian, Tyler R. Kartzinel, TM Palmer, Jacob R. Goheen, David Augustine, Mahesh Sankaran, CE Tarnita, and RM Pringle. “Large Herbivores Suppress Liana Infestation in an African Savanna.” PNAS 10.1073/pnas.2101676118 (2021): n. pag.
Kawakatsu, Mari, Yphtach Lelkes, Simon A. Levin, and CE Tarnita. “Interindividual Cooperation Mediated by Partisanship Complicates Madison’s Cure for ‘mischiefs of faction’.” PNAS 118 (2021): e2102148118.
Tokita, Christopher K., Guess, and CE Tarnita. “Polarized Information Ecosystems Can Reorganize Social Networks via Information Cascades.” PNAS 118 (2021): e2102147118.
Ulrich, Yuko, Mari Kawakatsu, Christopher Tokita, Jonathan Saragosti, Vikram Chandra, CE Tarnita, and Daniel J.C. Kronauer. “Response Thresholds Alone Cannot Explain Empirical Patterns of Division of Labor in Social Insects.” PLoS Biology 19.6 (2021): e3001269.


Baker, CCM, Jessica Castillo Vardaro, DF Doak, Johan Pansu, Jeremy Puissant, RM Pringle, and CE Tarnita. “Spatial Patterning of Soil Microbial Communities Created by Fungus-Farming Termites.” Molecular Ecology 29 (2020): 4487–4501.
Pourtois, Julie, Corina E. Tarnita, and Juan A. Bonachela. “Impact of Lytic Phages on Phosphorus- Versus Nitrogen-Limited Marine Microbes.” Frontiers in Microbiology (Aquatic Microbiology) (2020): n. pag. Print.
Rossine, Fernando, Ricardo Martinez-Garcia, Allyson Sgro, Thomas Gregor, and Corina E. Tarnita. “Eco-Evolutionary Significance of ’loners’.” PLoS Biology (2020): n. pag. Print.
Staps, Merlijn, and CE Tarnita. “How Geometry Shapes Division of Labor.” eLife 9:e63328 (2020): n. pag. Print.


Atkins, Justine, Ryan Long, Johan Pansu, Corina E. Tarnita, and Robert Pringle. “Cascading Impacts of Large-Carnivore Extirpation in an African Ecosystem.” Science 364 (2019): 173–177.
Stalmans, Marc, Tara Massad, Mike J. S. Peel, Corina E. Tarnita, and Robert Pringle. “War-Induced Collapse and Asymmetric Recovery of Large-Mammal Populations in Gorongosa National Park, Mozambique.” PLoS ONE 14.3 (2019): e0212864.
Staps, Merlijn, Jordi Gestel, and Corina E. Tarnita. “Emergence of Diverse Life Cycles and Life Histories at the Origin of Multicellularity.” Nature Ecology & Evolution (2019): n. pag.


Tarnita, Corina E. “Fast Evolution Unlocks Forbidden Communities.” Nature Ecology & Evolution 2 (2018): 1525–1526.
Tarnita, Corina. “Termite Patterning at Multiple Scales.” Current Biology 28 (2018): R1382–R1405.
Ulrich, Yuko, Jonathan Saragosti, Christopher Tokita, Corina E. Tarnita, and Daniel J.C. Kronauer. “Fitness Benefits and Emergent Division of Labour at the Onset of Group Living.” Nature 560 (2018): 635–638.


Gestel, Jordi, and Corina E. Tarnita. “On the Origin of Biological Construction, With a Focus on Multicellularity.” Proc Natl Acad Sci 114.42 (2017): 11018–11026.
Martinez-Garcia, Ricardo, and Corina E. Tarnita. “Seasonality Can Induce Coexistence of Multiple Bet-Hedging Strategies in Dictyostelium Discoideum via Storage Effect.” Journal of Theoretical Biology 426 (2017): 104–116.
Tarnita, Corina E. “The Ecology and Evolution of Social Behavior in Microbes.” Journal of Experimental Biology. Special Issue: Evolution of Social Behavior 220.1 (2017): 18–24.
Tarnita, Corina E., Juan A. Bonachela, Efrat Sheffer, Jennifer A. Guyton, Tyler C. Coverdale, Ryan A. Long, and Robert M. Pringle. “A Theoretical Foundation for Multi-Scale Regular Vegetation Patterns.” Nature 541 (2017): 398–401.
Thutupalli, Uppaluri, George W.A. Constable, Simon A. Levin, Howard A. Stone, CE Tarnita, and C.P. Brangwynne. “Farming and Public Goods Production in C. Elegans Populations.” Proc Natl Acad Sci 114.9 (2017): 2289–2294.


Constable, George W.A., Tim Rogers, Alan McKane, and Corina E. Tarnita. “Demographic Noise Can Reverse the Direction of Deterministic Selection.” Proc Natl Acad Sci 113.32 (2016): E4745–E4754. Print.
Martinez-Garcia, Ricardo, and Corina E. Tarnita. “Lack of Ecological and Life History Context Can Create the Illusion of Social Interactions in Dictyostelium Discoideum.” PLoS Computational Biology 12.12 (2016): e1005246.
Tarnita, Corina E. “Mathematical Approaches or Agent-Based Methods? Comment on ‘Evolutionary Game Theory Using Agent-Based Methods’ by Christoph Adami Et Al.” Physics of Life Reviews 10.1016/j.plrev.2016.10.017.(invited comment) (2016): n. pag. Print.
Widder, RJ Allen, Pfeiffer T, TP Curtis, Wiuf, WT Sloan, OX Cordero, Brown, Momeni, Shou, Kettle, HJ Flint, AF Haas, Laroche, JU Kreft, PB Rainey, Freilich, Schuster, Milferstedt, JR Meer, Grosskopf, Huisman, Free, Picioreanu, Quince, Klapper, Labarthe, BF Smets, Wang, Isaac Newton Institute Fellows (including CET), and OS Soyer. “Challenges in Microbial Ecology: Building Predictive Understanding of Community Function and Dynamics.” ISME J (2016): 1–12. Print.


Bonachela, Juan, Robert Pringle, Efrat Sheffer, Tyler Coverdale, Jen Guyton, Kelly Caylor, Simon Levin, and Corina Tarnita. “Termite Mounds Can Increase the Robustness of Drylands Ecosystems to Climatic Change.” Science 347 (2015): 651–655. Print.
Tarnita, Corina E., Alex Wahburne, Ricardo Martinez-Garcia, Allyson Sgro, and Simon Levin. “Fitness Tradeoffs Between Spores and Nonaggregating Cells Can Explain the Coexistence of Diverse Genotypes in Cellular Slime Molds.” Proc Natl Acad Sci 112 (2015): 2776–2781. Print.
Tarnita, Corina. “Fairness and Trust in Structured Populations.” GAMES (Special issue: Cooperation, Trust, and Reciprocity) 6 (2015): 214–230. Print.


Allen, and CE Tarnita. “Measures of Success in a Class of Evolutionary Models With Fixed Population Size and Structure..” Journal of Mathematical Biology 68 (2014): 109–143. Print.
Pringle, RM, JR Goheen, TM Palmer, GK Charles, DeFranco, Hohbein, AT Ford, and CE Tarnita. “Low Functional Redundancy Among Mammalian Browsers in Regulating an Encroaching Shrub (Solanum Campylacanthum) in African Savannah.” Proceedings of the Royal Society, B 281.1785 (2014): n. pag. Print.
Tarnita, Corina, and Peter Taylor. “Measures of Relative Fitness of Social Behaviors in Finite Structured Population Models.” American Naturalist 184.4 (2014): 477–488. Print.
Tarnita, Corina, Todd Palmer, and Robert Pringle. “Colonization and Competition Dynamics Can Explain Incomplete Sterilization Parasitism in Ant-Plant Symbioses.” Ecology Letters 17.10 (2014): 1290–1298. Print.


Rand, DG*, CE Tarnita*, Ohtsuki, and MA Nowak. “The Evolution of Fairness in the One-Shot Anonymous Ultimatum Game..” Proceedings of the National Academy of Sciences 110 (2013): 2581–2586 *joint first authors. Print.
Tarnita, CE, CH Taubes, and MA Nowak. “Evolutionary Construction by Staying Together and Coming Together..” Journal of Theoretical Biology 320 (2013): 10–22. Print.


Allen, Traulsen, CE Tarnita, and MA Nowak. “How Mutation Affects Evolutionary Games on Graphs..” Journal of Theoretical Biology 299 (2012): 97–105. Print.
Cavaliere, Sedwards, CE Tarnita, MA Nowak, and Csikasz-Nagy. “Prosperity Is Associated With Instability in Dynamical Networks..” Journal of Theoretical Biology 299 (2012): 126–138. Print.
Fu, CE Tarnita*, NA Christakis, Wang, DG Rand, and MA Nowak. “Evolution of in-Group Favoritism..” Nature Scientific Reports doi:10.1038/srep00460 (2012): *joint first authors. Print.
Gestel, MA Nowak, and CE Tarnita. “Evolution of Cell-to-Cell Communication in a Sporulating Bacterium..” PLoS Comp Biol 8(12): e1002818 (2012): n. pag. Print.


Tarnita, CE, Wage, and MA Nowak. “Multiple Strategies in Structured Populations..” Proceedings of the National Academy of Sciences 108 (2011): 2334–2337. Print.


Nowak, MA, CE Tarnita, and Antal. “Evolutionary Dynamics in Structured Populations..” Phil Trans R Soc B 365 (2010): 19–30. Print.
Nowak, MA, CE Tarnita, and EO Wilson. “Evolution of Eusociality..” Nature 466 (2010): 1057–1062. Print.
Nowak, MA, CE Tarnita, and EO Wilson. “Nowak Et Al. Reply..” Nature 471 (2010): E9-E10. Print.


Grigorov, Jorza, Patrikis, Stein, and CE Tarnita. “Verification of the Birch and Swinnerton-Dyer Conjecture for Specific Elliptic Curves..” AMS Journal "Mathematics of Computation" 78 (2009): 2397–2425. Print.
Nathanson, CG*, CE Tarnita*, and MA Nowak. “Calculating Evolutionary Dynamics in Structured Populations..” PLoS Comp Biol 5(12): e1000615 (2009): *joint first authors. Print.
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